Skip to main content



Millon, E.M., Haddad, A.E., Han Yan M. Chang, M.S., Najafizadeh, L., Shors, T.J. (2021). Human neural responses associated with the subjective passing of time. Submitted.

Millon, E.M., Lehrer, P. & Shors, T.J. (2021). Mental and Physical (MAP) Training with meditation and aerobic exercise enhances mental health and cognition in women living with HIV. Submitted.

Millon, E.M. & Shors, T.J. (2021). How mental health relates to everyday stress, rumination, trauma and interoception in women living with HIV: A factor analytic study, Learning and Motivation 73, 101680.

Lavadera, P., Millon, E.M., Shors, T.J. (2020). MAP Train My Brain: Meditation combined with aerobic exercise reduces stress and rumination while enhancing quality of life in medical students. The Journal of Alternative and Complementary Medicine. 26, 418-423.

Million E.M. and Shors T.J. Shors, T. J. (2019). Taking neurogenesis out of the lab and into the world with MAP Train My Brain, Behavioral Brain Research, 376, 112154 [Link].

Shors T.J., Chang H.Y. and Millon E.M. (2018). MAP Training My BrainTM: Meditation plus aerobic exercise lessens trauma in women with sexual violence more than either activity alone. Frontiers in Neuroscience, 12, 211. (More than 11K views and in the top 5% of all outputs scored by Altmetric).  [Link]

Millon E.M., Chang H.Y. and Shors T.J. (2018) Stressful life memories relate to ruminative thoughts in women with sexual violence history, irrespective of PTSD. Frontiers in Psychiatry 9:31 (More than 11K views and in the top 5% of all outputs).

Shors, T. J., Millon, E. M., Chang, H. Y. M., Olson, R. L., & Alderman, B. L. (2017). Do sex differences in rumination explain sex differences in depression?. Journal of Neuroscience Research, 95(1-2), 711-718. [Link] [PDF]

Difeo, G., Shors, T. J. (2017). Mental and physical skill training increases neurogenesis via cell survival in the adolescent hippocampus. Brain Research, 1654(Pt B), 95-101. [Link] [PDF]

Shors, T. J., & Millon, E. M. (2016). Sexual trauma and the female brain. Frontiers in Neuroendocrinology, 41, 87-98. [Link] [PDF]

Alderman B.L., Olson R.L., Brush C.J. and Shors T.J. (2016). Mental and Physical (MAP) Training: Combining meditation and aerobic exercise reduces depression and rumination while enhancing synchronized brain activity. Transl. Psychiatry 6, e726; doi:10.1038/tp.2015.225 [Last year, this publication had their highest impact via Altmetric and was their most read; it is still in top 5% with more than 11K views] [Link]

Shors, T. J. (2016). A trip down memory lane about sex differences in the brain. Philosophical Transactions of the Royal Society B, 371. doi: 10.1098/rstb.2015.0124 [Link] [PDF]

Shors, T. J., Tobόn, K., DiFeo, G., Durham, D. M., & Chang, H. Y. M. (2016). Sexual Conspecific Aggressive Response (SCAR): A Model of Sexual Trauma that Disrupts Maternal Learning and Plasticity in the Female Brain. Scientific Reports, 6. doi:10.1038/srep18960 [Link]

Alderman, B. L., Olson, R. L., Bates, M. E., Selby, E. A., Buckman, J. F., Brush, C. J., … & Shors, T. J. (2015). Rumination in major depressive disorder is associated with impaired neural activation during conflict monitoring. Frontiers in Human Neuroscience, 9, 269. [Link] [PDF]

Shors, T. J., Olson, R. L., Bates, M. E., Selby, E. A., & Alderman, B. L. (2014). Mental and Physical (MAP) Training: A neurogenesis-inspired intervention that enhances health in humans. Neurobiology of learning and memory, 115, 3-9. [PDF]

DiFeo, G., Curlik, D. M., & Shors, T. J. (2014). The motirod: a novel physical skill task that enhances motivation to learn and thereby increases neurogenesis especially in the female hippocampus. Brain research, 1621, 187-196. [PDF]

Shors, T. J. (2014). The adult brain makes new neurons, and effortful learning keeps them alive. Current Directions in Psychological Science, 23, 311-318. [PDF]

Curlik, D.M., DiFeo, G, Shors, T.J. (2014). Preparing for adulthood: Thousands upon thousands of new cells are born in the hippocampus during puberty, and most survive with effortful learning. Frontiers in Neuroscience, 8: 70. [PDF]

Maeng, L. Y., & Shors, T. J. (2013). The stressed female brain: neuronal activity in the prelimbic but not infralimbic region of the medial prefrontal cortex suppresses learning after acute stress. Frontiers in neural circuits, 7, 198. [PDF]

Curlik, D.M., Shors, T.J. (2013). Training your brain: Do mental and physical (MAP) training enhance cognition through the process of neurogenesis in the hippocampus? Neuropharmacology, 64, 506-514. [PDF]

Curlik, D.M., Maeng, L.Y., Agarwal, P.R., Shors, T.J. (2013). Physical Skill Training Increases the Number of Surviving New Cells in the Adult Hippocampus. PLoS One, 8(2): e55850. [PDF]

Anderson, M.L., Nokia, M.S., Govindaraju, K.P., Shors, T.J. (2012). Moderate drinking? Alcohol consumption significantly decreases neurogenesis in the adult hippocampus. Neuroscience, 224, 202-9. [PDF]

Nokia, M.S., Sisti, H.M., Choksi, M.R., Shors, T.J. (2012). Learning to Learn: Theta Oscillations Predict New Learning, which Enhances Related Learning and Neurogenesis. PLoS One, 2012;7(2):e31375 [PDF]

Shors T.J., Anderson, M., Curlik D.M., Nokia, M. (2012). Use it or lose it: How neurogenesis keeps the brain fit for learning. Behavioral Brain Research, special issue on neurogenesis, 227(2):450-8 [PDF]

Anderson, M.L. & Shors, T.J. Associative learning increases neurogenesis during a critical period. European Journal of Neuroscience, 33, 175-181, 2011. [PDF]

Waddell, J., Anderson, M.L., & Shors, T.J. Changing the rate and hippocampal dependence of trace eyeblink conditioning: slow learning enhances survival of new neurons. Neurobiology of Learning and Memory, 95, 159-165, 2011. [PDF]

Curlik D. & Shors T.J. (2011). Learning increases the survival of newborn neurons provided that learning is difficult to achieve and successful. Journal of Cognitive Neuroscience, 23(9), 2159-70. [PDF]

Shors T.J. Saving new brain cells. Scientific American, March Issue, 300, 46-52, 2009. [PDF]

Shors T.J. From stem cells to grandmother cells: How neurogenesis relates to processes of learning and memory. Cell Stem Cell, 3, 253-258, 2008. [PDF]

Dalla C., Papachristos E., Whetstone A.S., Shors T.J. Females learn trace memories better than males, thereby preserving more neurons in their hippocampus. Proceedings of the National Academy of Sciences, 106, 2927-2932, 2009. [PDF]

Shors T.J. Memory traces of trace memories: neurogenesis, synaptogenesis and awareness. Trends in Neuroscience, 27, 250-256, 2004. [PDF]

Leuner B., Falduto J., Shors, T.J. Associative memory formation increases the observation of dendritic spines in the hippocampus. Journal of Neuroscience, 23, 659-665, 2003. [PDF]

Shors T.J., Townsend D.A., Zhao M., Kozorovitskiy Y., Gould E. Neurogenesis may relate to some by not all types of hippocampal-dependent learning. Hippocampus, 12, 578-584, 2002. [PDF]

Shors T.J., Miesegeas G., Beylin A., Zhao M., Riedel T., Gould E. Neurogenesis in the adult is involved in the formation of trace memories. Nature, 410, 372-376, 2001. [PDF]

Gould E., Beylin A., Tanapat P., Reeves A., and Shors T.J. Learning enhances adult neurogenesis in the hippocampal region. Nature Neuroscience, 2, 260-265, 1999. [PDF]

Shors T.J. and Matzel L.D. Long-term potentiation (LTP): what’s learning got to do with it? Behavioral and Brain Sciences 20 (4), 597-655, 1997. [PDF]